ALT A two-panel diagram comparing Complex Environments and Simple Environments. In the complex panel, a jagged multicolored line graph sits beside a line drawing of a bacterium surrounded by colorful circles, triangles, and other shapes. In the simple panel, a flat pink line graph sits beside a line drawing of a bacterium surrounded only by identical pink circles. Below each panel, an arrow labeled 'Evolution' points to Controllability up, Thermodynamic Cost up for the complex panel, and Controllability down, Thermodynamic Cost down for the simple panel. At the bottom, a pink gear represents ATP and a blue gear represents GTP. The two gears are similarly sized in the complex panel, with one gear much larger than the other in the simple panel.